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| Keys
  to the Hymenoptera of European Russia(Email Contacts)    [The following keys are in a form commonly
  used in Eastern Europe.  If the
  statement is true,  proceed to the next line, whereas if it is
  false, go to the line shown in parentheses within a specific Superfamily]             (Note:  Not all illustrations are European species):        ------
  Please CLICK for specific Superfamily:   Chrysidoidea    Chalcidoidea   
  Proctotrupoidea   Ceraphronoidea     Keys to the Hymenoptera of European U.S.S.R.-- Chrysidoidea (=
  Bethyloidea)   (Derived from
  Alekseev, 1978, Boucek, 1951, 1956; Debauche, 1948, Ferriere, 1947, Graham, 1969,
  Hedquist,1975, Kieffer, 1914, Kozlov, 1978, Masner, 1976, Masner &
  Dessart, 1967, Medvedev, 1978, Nikolskaya,1952, Nikolskaya & Zerova,
  1978, Peck et al., 1964, Richards, 1939, Tryapitcyn, 1978 by E. F. Legner)   =  =  = 
  =  =  =  =  = 
  =  =  =  =  =  =  =  = 
  =  =  =   1 (2).    Antennae with 12-13 segments, attached near mouth margin; body
  long and narrow; usually somewhat flat insects .......... Bethylidae   <Habits>; <Adults>
  & <Juveniles>   2 (1).    Antennae with 10 segments,
  attached at projection near dorsal margin of clypeus   3 (4).    Head of female not pear-shaped; fore
  tarsi female with claws; if claws absent, then female dipterous (genus Aphelopus); fore wings without closed discoidal
  cell  ........... Dryinidae   <Habits>; <Adults>
  & <Juveniles>   4 (3).    Head of female pear-shaped; fore tarsi
  of female without claws; female always wingless; fore wings with closed
  discoidal cell  .......... Embolemidae   <Habits>; <Adults>        Keys to the Hymenoptera of European U.S.S.R.-- Chalcidoidea   (Derived from Medvedev, 1978 by E. F. Legner)     1 (76).    Wings not reduced; fore wings, when
  folded back, extend beyond tip of abdomen.   2 (3).    Tarsi with 3 segments; very minute species (body length 0.30 - 1.40 mm.); antennae usually very short, with 4-8 segments; hairs on fore wings often arranged
  in longitudinal rows; endoparasitoids of insect eggs .......... Trichogrammatidae   <Habits>; <Adults>
  & <Juveniiles>   3 (2).    Tarsi with 4-5 segments   4 (21).    Tarsi of at least middle pair of legs with 4 segments   5 (8).    Middle tarsi with 4 segments; fore- and hind tarsi with 5 segments   6 (7).    Antennae with 12 segments; fore wings with fairly long
  postmarginal and radial veins; body length 2.0 - 3.5 mm
  .......... Pteromalidae  (only
  females of Macromesinae)   <Habits>; <Adults>
  & <Juveniles>   7 (6).    Antennae with 8 segments; fore wings without postmarginal vein
  and radial vein short; body length  0.5 - 0.8 mm .......... Aphelinidae  (some
  species of Trichaporus & Encarsia)  <Habits>; <Adults>
  & <Juveniles>   8 (5).    Tarsi of all legs with 4 segments   9 (10).    Antennal sockets closer to inner margin of eyes than to each other;
  distance from antennal socket to eye margin less than diameter of antennal
  socket; frons above antennal sockets with straight, deep, transverse stria;
  two pairs of longitudinal striae originate from ends of transverse stria and
  continue along inner margin of eyes; antennae without rings, proceed directly
  from pedicel; marginal vein of fore wings short; radial vein rudimentary, its
  apex in most cases falling short of mid wing; base of hind wings represented
  by only submarginal vein, and speculum does not reach wing base; margins of
  hind wings usually almost parallel beyond hamuli (= series of hooks); body
  not metallic; endoparasitoids of insect eggs ......... Mymaridae   <Habits>; <Adults>
  & <Juveniles>   10 (9).    Antennal sockets usually not closer to inner margin of eyes than to
  each other (if closer to inner margin of eyes, then frons without deep
  transverse stria and longitudinal striae originating from its ends; if
  transverse frontal stria present, usually not straight)   11 (12).    Marginal vein of fore wings dotted, or almost dotted; sides of
  mesothorax not delineated; fore wings either with very long marginal fimbria,
  not shorter than maximum width of wing (Anthemus), or antennae of female with 5 segments, 2 rings, and very small, nonsegmented clava  (Arrhenophagus); body length 0.50 - 0.70 mm
  ........ Encyrtidae  (only Anthemus & Arrhenophagus)   <Habits>; <Adults-1>
  & <Adults-2>; & <Juveniles>   12 (11).    Marginal vein of fore wings much longer than wide; sides of
  mesothorax delineated   13 (14).    Antennal funicle with 6 segments (excluding very thin rings between pedicel and 1st segment of funicle); fore wings without speculum (= area
  running obliquely from parastigma to hind margin) (base of wing pubescent)
  ......... Tetracampidae  (only
  males of Tetracampinae)  <Habits>; <Adults-1>
  & <Adults-2>; & <Juveniles>   14 (13).    Antennal funicle, at most, with 5 segments
  (excluding slender rings between pedicel and 1st segment
  of funicle, which may vary in number from 1-5); fore wings usually with speculum   15 (16).    Hind coxae highly disk-like, broad and flat, almost lamellate; outer surface
  of tibiae with numerous short, dark, coarse bristles arranged in somewhat
  zigzag pattern; fore wings narrow, with almost parallel anterior and
  posterior margins; marginal vein of fore wings very long ......... Elasmidae  (only Elasmus)   <Habits>; <Adults>
  & <Juveniles>   16 (15).    Hind coxae broad, disk-like; outer surface of tibiae without zigzag
  pattern or dark bristles   17 (18).    Postmarginal vein of fore wings equal in length to radial vein, or
  longer; radial vein not very short ........... Eulophidae   <Habits>; <Adults>
  & <Juveniles>   18 (17).    Postmarginal vein of fore wings either absent or distinctly shorter
  than radial vein; radial vein of fore wings very short or even rudimentary   19 (20).    Abdomen distinctly attenuates at juncture with median segment;
  abdominal petiole always well defined, even if sometimes represented by only
  a transverse ring; endophragm usually does not continue into abdomen; angle
  between radial vein of fore wings and wing margin not less than 35-deg.; body often (at least partly) metallic; if not metallic,
  then scutellum with two deep longitudinal striae near midpoint .......... Eulophidae   <Habits>; <Adults>
  & <Juveniles>   20 (19).    Abdomen appears almost sessile, i.e., its base
  almost equal in width to median segment; abdominal petiole very difficult to distinguish;
  endophragm continues into abdomen; angle between radial vein of fore wings
  and wing margin usually very acute; body not metallic; scutellum without deep
  median longitudinal striae .......... Aphelinidae   21 (4).    Tarsi of all legs with 5 segments   22 (23).    Antennal sockets closer to inner margin
  of eyes than to each other; distance from antennal socket to margin of eye
  less than diameter of antennal socket; frons above antennal socket with
  straight, deep, transverse stria; two pairs of longitudinal striae originate
  from ends of transverse stria and continue along inner margin of eyes;
  antennae without rings, proceed directly from pedicel; marginal vein of fore
  wings short; radial vein rudimentary, its apex in most cases falling short of
  mid wing; base of hind wings represented by only submarginal vein, and
  speculum does not reach wing base; margins of hind wings usually almost
  parallel beyond hamuli; body not metallic; endoparasitoids of insect eggs
  .........  Mymaridae   <Habits>; <Adults>
  & <Juveniles>   23 (22).    Antennal sockets usually not closer to inner margin of eyes than to each
  other (if closer to inner margin of eyes, then frons without transverse deep
  stria and longitudinal striae originating from its ends; if transverse
  frontal stria present, usually not straight)   24 (25).    Middle coxae situated almost at midpoint of episternum of mesothorax,
  sometimes even before its midpoint; mesothorax very short; sides of
  mesothorax convex, consist of highly enlarged episternum; spurs of middle
  tibiae usually highly thickened and long; 1st segment
  of middle tarsi usually much longer than other segments ); shield of
  mesonotum usually without parapsidal grooves (if such present, very fine and
  shallow); cerci usually situated some distance from tip of abdomen  ......... 
  Encyrtidae   <Habits>; <Adults-1>
  & <Adults-2>; & <Juveniles>   25 (24).    Middle coxae situated behind level of midpoint of episternum of
  mesothorax; sides of mesothorax usually delineated, not convex, and do not
  broaden  (except in && of
  Eupelmidae; spurs of middle tibiae usually not very long and thick (except in
  Eupelmidae & Signiphoridae)   26 (27).    Middle tarsi thickened at base and attenuate
  toward apex; 1st segment of middle tarsi, and usually some
  subsequent segments, with 2 rows of short, thick spinules on lower
  surface; middle coxae separated from trochanter lobes of mesothorax by
  membrane and, as a result, coxae able to move straight forward; spurs of
  middle tibiae very long and thick, but without spines; sides of mesothorax
  highly enlarged, almost not demarcated, but set off from mesothorax by
  longitudinal suture; pronotum often longitudinally divided medially by
  groove, membranous stria, or carina; antennae mostly with one ring, 7-segmented funicle, and nonsegmented  or 3-segmented clava (also see .......... Eupelmidae  (only
  females)   <Habits>; <Adults>
  & <Juveniles>   27 (26).    Middle tarsi not thickened at apex [sic]
  and without row of thick spinules on lower surface; middle coxae not
  separated from mesothorax by membrane and unable to move straight forward;
  spurs of middle tibiae not elongated and not thickened but, if sometimes very
  long, then with long spines (Signiphoridae); sides of mesothorax usually
  separated by suture, which extends from base of middle coxae toward base of
  fore wings; pronotum usually not medially divided by longitudinal groove or
  carina.   28 (31).    Postspiracular sclerite situated in same plane
  as lateral part of pronotum or fused with it; pronotum firmly attached to
  episternum of mesothorax; thorax (in profile) short and high; 3rd and 4th abdominal tergites highly enlarged and
  usually occupy entire or almost entire dorsal part of abdomen   29 (30).    Pronotum not visible in dorsal view, concealed
  by highly convex or forwardly protruding shield of mesonotum; abdominal
  petiole long, often slightly longer than wide; radial vein of fore wings
  forms a right angle or almost a right angle with wing margin; head (facial
  view) usually somewhat triangular; mandibles almost always falciform
  (sometimes very short, straight, arranged vertically .......... Eucharitidae   <Habits>; <Adults>
  & <Juveniles>   30 (29).    Mesonotum distinctly visible in dorsal view
  (except in Philomides Haliday);
  abdominal petiole very short, transverse; radial vein of fore wings forms an acute
  angle with wing margin; head (facial view) not triangular; mandibles not
  falciform .......... Perilampidae   <Habits>; <Adults>
  & <Juveniles>   31 (28).    Postspiracular sclerite not situated in same
  plane as lateral part of pronotum and not fused with it   32 (35).    Species associated with fig fruits (Ficus
  carica).   33 (34).    Length of raised part of ovipositor constitutes about 1/4 length of abdomen; ovipositor not overlapped
  by last abdominal tergite on dorsal side; fore wings 2X longer than their maximum width; 1st segment of antennal funicle with process; mandibular base with
  process with transverse carinae .......... Agaonidae  (only
  females)   <Habits>; <Adults>    34 (33).    Ovipositor
  very long, much longer than abdomen; about 1/3rd of ovipositor overlapped by last abdominal tergite on dorsal
  side; fore wings 3X longer than max. width; mandibles with 2 teeth .......... Torymidae  (only
  females of Idarninae)   <Habits>; <Adults>
  & <Juveniles>   35 (32).    Species not associated with fig fruits   36 (37).    Last (9th) abdominal tergite with notch on back side
  and small lobe (epipygium) in middle of this notch; cerci (= pygostyles)
  longer than wide; ovipositor usually long; hind coxae several times larger
  than fore coxae ......... Torymidae  (only
  females)   <Habits>; <Adults>
  & <Juveniles>   37 (36).    Last (9th)
  abdominal tergite without notch on back side and without lobe; cerci usually not
  longer than wide   38 (43).    Hind femora highly broadened and dilated, only 1.5 - 3.0 times longer than max. width, and ventral margin with teeth or
  serrated; hind tibiae usually distinctly curved   39 (40).    Postspiracular sclerite large, extending notably downward; distance
  from tegula to posterior margin of pronotum almost equal to tegula; body
  usually metallic; antennae usually attached at least slightly above level of
  lower margin of eyes; postmarginal vein of fore wings shorter than marginal
  vein .......... Torymidae   <Habits>; <Adults>
  & <Juveniles>   40 (39).    Postspiracular sclerite usually poorly perceptible; if sometimes
  large (in some members of Leucospidae), tegulaealmost touch pronotum; body usually
  not metallic.   41 (42).    Tegulae not more than 2X longer than wide, their anterior margin
  situated at a notable distance from margin of pronotum; distance between
  apices of axillae not greater than axillar width; parapsidal grooves usually
  continuous, sometimes shallow on back side; fore wings not folded along their
  length; ovipositor does not turn 
  toward dorsal side of abdomen .......... Chalcididae   <Habits>; <Adults>
  & <Juveniles>   42 (41).    Tegulae 2.5 - 3.5 times longer than wide, their anterior margin
  almost reaching margin of pronotum; axillae very small; distance between
  their apices several times greater than axillar width; axillae often
  indistinctly separated from scutellum; shield of mesonotum without parapsidal
  grooves; sometimes grooves barely perceptible on front; fore wings folded
  along length, as in wasps of the family Vespidae; ovipositor turns toward
  dorsal surface of abdomen .......... Leucospidae   <Habits>; <Adults>
  & <Juveniles>   43 (38).    Hind femora not highly broadened (if sometimes notably broadened,
  then ventral surface without teeth or denticles); hind tibiae do not curve or
  only slightly so.   44 (47).    Pronotum (except neck) long, almost rectangular, its length at least 2/3rds length of mesonotal shield; genae with
  acute margin; surface of head and dorsal side of thorax with very dense,
  almost fused, coarse punctation; body usually not metallic   45 (46).    Antennal
  funicle, at most, with 6 segments; inner margin of eyes diverges
  slightly toward lower side ......... Eurytomidae   <Habits>; <Adults>
  & <Juveniles>   46 (45).    Antennal funicle with 7 segments (excluding ring); inner margin of
  eyes diverges notable toward lower side ......... Pteromalidae   <Habits>; <Adults>
  & <Juveniles>   47 (44).    Pronotum transverse (if pronotum long and almost rectangular, then
  genae either without sharp margin, or head and dorsal part of thorax without
  coarse and deep punctation)   48 (63).    Pronotum (excluding neck) almost rectangular, long, its length
  constituting at least 1/2 length of
  mesonotal shield;mesonotal shield with continuous parapsidal grooves (=
  mesoscutal marks).   49 (50).    Marginal vein of fore wings 4-8X longer than radial vein (postmarginal vein not longer or barely
  longer than radial vein); hairs of abdominal cerci very long; antennal sockets
  either touch margin of mouth or face, and with  tubercles or crests .......... Pteromalidae   <Habits>; <Adults>
  & <Juveniles>   50 (49).    Marginal vein of fore wings either maximally 3X longer than radial vein, or hairs of cerci short; antennal
  sockets do not touch margin of mouth and face, and without tubercles or
  crests.   51 (52).    Antennae
  with 10-11 segments,
  of which 4-6 constitute
  funicle; sides of metathorax not distinguished  or indistinctly separated from median segment; body usually
  without metallic glaze ......... Eurytomidae   <Habits>; <Adults>
  & <Juveniles>   52 (51).    Antennae with 12-13 segments
  (usually 13, rarely 12), of which
  6-9 constitute
  funicle.   53 (54).    Body black, without metallic glaze; median
  segment with median longitudinal groove; scutellum without distinctly
  discernible frenum ......... Eurytomidae   <Habits>; <Adults>
  & <Juveniles>   54 (53).    Body at least partly with metallic
  glaze, or median segment without median longitudinal groove; scutellum
  usually with frenum distinguished by distinct, deep stria.   55 (56).    Abdomen with well-defined, almost rectangular,
  reticulate or striated petiole; anterior margin of sides of metathorax do not
  curve; hind femora not dilated and without ventral serration; radial vein of
  fore wings with poorly broadened apex ............ Pteromalidae   <Habits>; <Adults>
  & <Juveniles>   56 (55).    Abdomen with petiole relatively less visible, almost
  conical, and smooth or almost smooth   57 (58).    Antennae attached below level of lower margin
  of eyes; fore wings either virtually without speculum and marginal vein
  barely longer than radial, or vertex with 6-8 large, dark, and forwardly directed hairs .......... Pteromalidae   <Habits>; <Adults>
  & <Juveniles>   58 (57).    Antennae attached at level of lower margin
  of eyes or above margin.   59 (60).    Antennal funicle with 9 segments; radial vein of fore wings with
  poorly broadened apex ......... Pteromalidae   <Habits>; <Adults>
  & <Juveniles>   60 (59).    Antennal funicle, at most, with 7 segments   61 (62).    Neck of pronotum with sharp margin in front;
  abdomen shorter than thorax, almost spherical; ovipositor does notprotrude;
  mandibles large, with 2 teeth ......... Pteromalidae   <Habits>; <Adults>
  & <Juveniles>   62 (61).    Neck of pronotum without sharp margin in front .......... Torymidae   <Habits>; <Adults>
  & <Juveniles>   63 (48).    Pronotum (excluding neck) either not rectangular or shorter than 1/2 length of mesonotal shield   64 (65).    Hind tibiae with 2 strong apical spurs, of which one or both curve; hind coxae almost
  equal in length to hind femora, with slender carina along dorsal margin;
  middle segments of abdomen with unique sculpture; marginal  vein of fore wings 6-9X longer than very short radial vein; antennae
  with 13 segments ........ Ormyridae   <Habits>; <Adults>    65 (64).    Hind tibiae often with only one apical spur (if with 2, then both spurs straight); hind coxae usually shorter; middle
  segments of abdomen without unique sculpture.   66 (67).    Hind coxae highly broadened  and almost lamellate; hind tibiae with
  short row of 4-6 long
  bristles on dorsal margin and row of 4 such
  bristles on outer surface; apices of hind tibiae with 2 long spurs; fore wings with 1-3 individual bristles near apex of submarginal vein; antennae with
  9 segments ......... Elasmidae   <Habits>; <Adults>
  & <Juveniles>   67 (66).    Hind coxae neither broad nor flat; hind tibiae
  without such rows of long bristles; apices of hind tibiae often with only one
  spur.   68 (69).    Axillae indistinctly separated from scutellum,
  forming with it a single transverse sclerite; median segment with triangular
  middle area; abdomen sessile, its basal segment equal in width to median
  segment; antennae with 5-7 segments, 2-4 rings, and very long nonsegmented clava; fore
  wings with long marginal fimbria, equal in length to at least 1/3rd wing width; middle tibiae with long spur
  ........ Signiphoridae   <Habits>; <Adults>
  & <Juveniles>   69 (68).    Axillae distinctly separated from
  scutellum; median segment without triangular middle area; abdomen somewhat
  petiolate, or almost so; if abdomen appears sessile, then its basal segment
  at least slightly narrower than median segment.   70 (71).    Antennae with 3-9 segments; postmarginal vein of fore wings not
  longer than 0.16 length of
  marginal vein; abdomen almost sessile; abdominal petiole, if present, highly
  transverse (= at right angles to longitudinal axis) and difficult to discern;
  postphragm extends atleast slightly into abdomen; pronotum very short and
  highly transverse; scutellum at least slightly transverse .......... Aphelinidae   <Habits>; <Adults>
  & <Juveniles>   71 (70).    Antennae with 10-13 segments; postmarginal
  vein of fore wings usually well developed and equal in length to radial vein
  or longer.   72 (73).    Median segment pubescent; middle 1/3rd with some hairs so that only narrow median
  strip remains glabrous; spur of fore tibiae weak and straight; fore wings
  without speculum; pronotum not shorter than shield of mesonotum; scutellum
  with 4 long bristles ......... Tetracampidae  (only
  females of Tetracampinae)   <Habits>; <Adults>    73 (72).    Median segment pubescent at least in middle 1/3rd; spur of fore tibiae usually strong and
  curved; fore wings often with speculum; pronotum often comparatively short;
  scutellum usually with more than 4 bristles.   74 (75).    Females:  Spur
  of fore tibiae weak, almost straight, and almost equal in length to 1/4 of 1st segment
  of fore tarsi; antennae with one ring, 6-segmented
  funicle, and 3-segmented clava.  Males:  Fore
  wings with large black longitudinal dilation that occupies marginal vein and
  distal part of submarginal .  Females & Males:  Pronotum bell-shaped, its length ca. 3/4ths length of shield of mesonotum .......... Tetracampidae  (only Platynocheilus)   <Habits>; <Adults>    75 (74).    Females:  Spur
  of fore tibiae strong and long, almost always curved (if spur straight, then
  antennal formula different).  Males:  Fore wings without large black dilatation .......... Pteromalidae   <Habits>; <Adults>
  & <Juveniles>   76 (1).    Wings highly reduced or rudimentary.   77 (78).    Tarsi with 3 segments; very minute species (body length 0.3 - 1.3 mm.); antennae usually very short, with 4-8 segments; endoparasitoids of insect eggs
  .......... Trichogrammatidae   <Habits>; <Adults>
  & <Juveniiles>   78 (77).    Tarsi with 4-5 segments
  (only in males of Blastophaga, found in the ovaries of the fig Ficus
  carica, do fore
  tarsi consist of 2 segments)   79 (82).    Tarsi with 4 segments.   80 (81).    Antennal sockets closer to inner margin of eyes than to each
  other; distance from antennal socket to margin of eye less than diameter of
  antennal socket; antennae slender, without rings (also see couplet 9) .......... Mymaridae   <Habits>; <Adults>
  & <Juveniles>   81 (80).    Antennal sockets closer to each other than to
  inner margin of eyes .......... Eulophidae   <Habits>; <Adults> & <Juveniles>   82 (79).    Tarsi with 5 segments.   83 (86).    Found in the ovaries of the fig Ficus carica   84 (85).    Antennae with 3 segments;
  head broader than long ......... Agaonidae  (only
  males of Blastophaga)   85 (84).    Antennae with 11 segments;
  head almost squared ........... Torymidae  (only
  males of Philotrypesis)   <Habits>; <Adults> & <Juveniles>   88 (87).    Middle coxae situated near posterior end of mesepisternum   89 (90).    Middle tarsi thickened at base and attenuate toward apex; 1st segment of middle tarsi (and often some subsequent segments)
  on lower side with 2 rows of short, thick spinules;
  mesepisternum  highly enlarged,
  convex, without femoral groove, but separated from mesothorax by deep stria
  or suture; middle coxae with membranousparts on lower side near base
  ........... Eupelmidae   <Habits>; <Adults>
  & <Juveniles>   90 (89).    Middle tarsi not thickened at base and without rows of short, thick
  spinules on lower side; mesepisternum (in addition to a few members of
  Aphelinidae in which antennae with 7-8 segments) separated by femoral groove; middle tarsi without
  membranous parts on lower side near base   91 (92).    Antennae with 6-8 segments; pronotum not long, shorter than shield of mesonotum
  ........... Aphelinidae   <Habits>; <Adults>
  & <Juveniles>   92 (91).    Antennae wither with 11-13 segments, or pronotum longer than shield of
  mesonotum   93 (94).    Pronotum broadens toward front; body not metallic,
  and covered with long hairs; found in stems of wheat, phytophagous,. Eurytomidae  (only Philachyra)   <Habits>; <Adults>
  & <Juveniles>   94 (93).    Pronotum does not broaden toward front (see couplet 49) .......... Pteromalidae   <Habits>; <Adults>
  & <Juveniles>       Keys to the Hymenoptera of European U.S.S.R.-- Proctotrupoidea   (Derived from Kozlov, 1978 by E. F. Legner)     1 (4).    First (basal) antennal segment 1.5X longer than max. width; hind wings with anal lappet   2 (3).    Antennae with 16 segments; tarsal claws crestate; abdominal
  petiole at least 2X longer than wide; fore wings with 1M (1st discoidal cell); female ovipositor not
  protruding from abdominal valves; parasitoids of larvae of Chrysopidae
  ......... Heloridae   <Habits>; <Adults>
  & <Juveniles>   3 (2).    Antennae
  with 13 segments; tarsal claws simple, rarely with 2-3 denticles; abdominal petiole not longer than
  max. width, at least not in dorsal view; fore wings without 1 M; female ovipositor protrudes from abdominal valves;
  parasitoids predominantly of larvae of beetles, rarely larvae of Diptera
  .......... Proctotrupidae   <Habits>; <Adults>
  & <Juveniles>   4 (1).    First (basal) antennal segment at least 3X (usually more than 3X) longer
  than max. width; hind wings without anal appet    5 (10).    Abdominal petiole consists of one segment   6 (7).    Tergites and sternites of abdominal petiole
  fused; frons with process; antennae attached at frontal projection at level
  of midpoint of eye; if attachment different, then antennae with 14-15 segments; abdomen roundish on sides, without
  sharp margin; parasitoids of larvae and pupae of Diptera, rarely of larvae of
  bethyloid wasps ........... Diapriidae   <Habits>; <Adults>
  & <Juveniles>   7 (6).    Tergites and sternites of abdominal petiole not fused; frons without
  process; antennae attached almost at level of lower margin of eyes near
  opening of mouth; abdomen bordered on sides or with sharp margin.   8 (9).    Antennae usually with 12 segments, rarely 11 segments; if exceptionally with 7 or 10 segments, then fore wingsat least with marginal vein, or 7th antennal segment forms very flat clava; parasitoids of insect
  & spider eggs  ............. Scelionidae   <Habits>; <Adults>
  & <Juveniles>   9 (8).    Antennae
  usually with 10 segments; rarely with 7-9; fore wings with only subcostal vein, often without
  veins; parasitoids of gall midges, whiteflies, coccids, and rarely eggs of
  beetles .......... Platygastridae   <Habits>; <Adults>
  & <Juveniles>   10 (5).   
  Abdominal petiole consists of 2 segments;
  fore wings like Mymaridae (long processes) .......... Mymarommatidae (normally placed in Chalcidoidea)   <Habits>; <Adults>      Keys to the Hymenoptera of European U.S.S.R-- Ceraphronoidea <Overivew>   (Derived from Alekseev, 1978 by E. F. Legner)   1 (2).    All tibiae with 2 spurs at apex; longer spur of fore tibiae
  apically bifurcate; antennae with 11 segments
  in both sexes; antennal funicle of females fusiform, thick in middle;
  mesonotum with parapsidal and median grooves over greater part; pterostigma
  usually semielliptical, rarely linear; anterior margin of 1st abdominal tergite elongated toward front and constricts into a
  "neck"; Waterston organ 
  absent ........... Megaspilidae   <Habits>; <Adults>    2 (1).   
  Middle tibiae with one spur at apex; other tibiae with 2; longer spur on fore tibiae not bifurcate; antennae of females
  with 9-10 segments;
  antennae of males with 11 segments; antennal funicle of females with
  somewhat distinct clava; mesonotum with only median groove or all grooves
  entirely absent; pterostigma linear; anterior margin of 1st abdominal tergite does not constrict and bordered by
  transverse suture; Waterston organ present 
  ...... Ceraphronidae   <Habits>; <Adults>
  & <Juveniles>   3 (--).  Not included in this key is a small family with
  hosts in the Diptera, Hymenoptera, Lepidoptera, Coleoptera and
  Homoptera.  They are more generally
  found as secondary external parasitoids of Aphididae and dactylopine
  Coccidae, through Braconidae and chalcidoid primaries.  Calliceras abnormis Perk. attacks mature dryinid larvae in their
  cocoons in Hawaii, and Lygocerus semiramosus Kieff, is a secondary parasitoid of Chermidae through Encyrtidae
  in India (Clausen 1940/1962)
  ........... Calliceratidae    <Habits>; <Adults>    = = = = = = = = = = = = = = = = = = = =   REFERENCES:   Alekseev,
  V. N.  1978a.  Superfamily Ceraphronoidea (Ceraphronids).
  p. 1213-.  In:  G. S. Medvedev (ed.) 1987,       Keys to the Insects of the European
  Part of the USSR. Vol. 3 Hymenoptera, Pt. 2.  Akad. Nauk., Zool.
  Inst.,       Leningrad, SSSR. (trans. fr. Russian,
  Amerind. Publ. Co., Pvt. Ltd., New Delhi). 
  1341 p   Boucek,
  Z.  1951.  The first revision of the European species
  of the family Chalcidae (Hymenoptera). 
  Acta. Ent.       Mus. Nat. Pragae 17, Suppl. 1.  108 p.   Boucek,
  Z.  1956.  Notes on the Czechoslovak Perilampidae
  (Hymenoptera- Chalcidoidea).  Acta.
  Faun. Ent. Mus.       Nat. Prague 1:  83-98.   Debauche,  1948.  Etude sur les Mymarommatidae et les
  Mymaridae de la Belgique (Hymenoptera, Chalcidoidea).        mem. Mus. Hist. Nat. Belg. 108:  42-6.   Ferrière,  1947.  Les espèces européenes du genre Elasmus
  Westw. (Hym., Chalc.).  Mitt. Schweiz.
  Ent. Ges. 29(6):      565-80.   Graham, M.
  W. R. de V.  1969.  The Pteromalidae of
  northwestern Europe (Hymenoptera: Chalcidoidea).  Bull.       Brit. Mus. Nat. Hist. Ent., Suppl. No. 16.  908 p.   Hedqvist,  1975.  Notes on Embolemidae and Bethylidae in
  Sweden with a description of a new genus and species       (Hym., Bethyloidea).  Ent. Tidschr. 96:  3-4, 121-32.   Kieffer,  1914.  Bethylidae.  Das Tierreich 41:  298-559.   Kozlov, M.
  A.  1978/1987.  Order Hymenoptera,
  Superfamily Proctotrupoidea (Proctotrupoids).  In:  G. S.       Medvedev (ed.) 1987, Keys to the Insects of the European Part of the USSR. Vol. 3 Hymenoptera, Pt. 2.  Akad.
        Nauk., Zool. Inst., Leningrad, SSSR.
  (trans. fr. Russian, Amerind. Publ. Co., Pvt. Ltd., New Delhi).  1341 p   Masner,  1976.  A revision of the Ismarinae of the New
  World (Hymenoptera, Proctotrupoidea, Diapriidae).  Canad.       Ent. 108:  1243-66.   Masner
  & Dessart.  1967.  La reclassification des
  categories taxonomiques superieures des Ceraphronoidea       (Hymenoptera).  Bull. Inst. Sci. Nat. Belg. 43(2):  1-33.   Medvedev,
  G. S. (ed.)  1978/1987.  Keys
  to the Insects of the European Part of the USSR. Vol. 3 Hymenoptera, Pt.       2.  Akad. Nauk., Zool. Inst., Leningrad, SSSR.
  (trans. fr. Russian, Amerind. Publ. Co., Pvt. Ltd., New Delhi).        1341 p.   Nikolskaya,
  M. N.  1952.  Chalcid Fauna of the Soviet Union
  (Chalcidoidea).  Key to the Fauna of
  the Soviet Union       44:  1-575.   Medvedev,
  G. S. (ed.)  1978/1987.  Keys
  to the Insects of the European Part of the USSR. Vol. 3 Hymenoptera, Pt.       2.  Akad. Nauk., Zool. Inst., Leningrad, SSSR.
  (trans. fr. Russian, Amerind. Publ. Co., Pvt. Ltd., New Delhi).         1341 p.   Medvedev,
  G. S.  1978/1987.  Order Hymenoptera,
  Superfamily Bethyloidea (Bethyloid Wasps). 
  In:  G. S.       Medvedev (ed.) 1987, Keys to the Insects of the European Part of the USSR. Vol. 3 Hymenoptera, Pt. 2.  Akad.
        Nauk., Zool. Inst., Leningrad, SSSR. (trans. fr. Russian,
  Amerind. Publ. Co., Pvt. Ltd., New Delhi). 
  1341 p.   Medvedev,
  G. S. (ed.)  1978/1987.  Keys
  to the Insects of the European Part of the USSR. Vol. 3 Hymenoptera, Pt.       2.  Akad. Nauk., Zool. Inst., Leningrad, SSSR.
  (trans. fr. Russian, Amerind. Publ. Co., Pvt. Ltd., New Delhi).        1341 p.   Medvedev,
  G. S.  1978/1987.  Order Hymenoptera,
  Superfamily Bethyloidea (Bethyloid Wasps). 
  In:  G. S.       Medvedev (ed.) 1987, Keys to the Insects of the European Part of the USSR. Vol. 3 Hymenoptera, Pt. 2.  Akad.
        Nauk., Zool. Inst., Leningrad, SSSR.
  (trans. fr. Russian, Amerind. Publ. Co., Pvt. Ltd., New Delhi).  1341 p.   Nikolskaya,
  M. N. & M. D. Zerova.  1978.  Superfamily Chalcidoidea,
  Family Torymidae, p. 651.  In:  G. S.       Medvedev (ed.) 1978/1987, Keys to the Insects of the European Part of
  the USSR. Vol. 3 Hymenoptera, Pt. 2.        Akad. Nauk., Zool. Inst., Leningrad,
  SSSR. (trans. fr. Russian, Amerind. Publ. Co., Pvt. Ltd., New Delhi).  1341       p.   Peck, O.,
  Z. Bou…ek & A. Hoffer.  1964.  Keys to the
  Chalcidoidea of Czechoslovakia (Insecta: Hymenoptera).        Mem. Ent. Soc. Canada 34:  120 p.   Richards,
  O. W.  1939.  The British Bethylidae (s.l.)
  (Hymenoptera).  Trans. Roy. Ent. Soc.,
  London 89(8):  297-      344.   Trjapitcyn,
  V. A.  1978a.  Superfamily Bethyloidea, Family
  Bethylidae, p. 3-.  In:  G. S. Medvedev (ed.) 1978/1987,       Keys to the Insects of the European
  Part of the USSR. Vol. 3 Hymenoptera, Pt. 2.  Akad. Nauk., Zool.
  Inst.,       Leningrad, SSSR. (trans. fr. Russian,
  Amerind. Publ. Co., Pvt. Ltd., New Delhi). 
  1341 p.   |